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1.
1) The Compositae in Tibet so far known comprise 508 species and 88 genera,
which nearly amounts to one fourth of the total number of genera and one third of the
total number of species of Compositae in all China, if the number of 2290 species and 220
genera have respectively been counted in all China. In Tibet there are all tribes of Com-
positae known in China, and surprisingly, the large tribes in Tibetan Compositae are
also large ones in all China and the small tribes in Tibet are also small ones in all China.
Generally speaking, the large genera in Tibet are also large ones in all China and the
small genera in Tibet are likewise small ones in all China. In this sense it is reasonable to
say that the Compositae flora of Tibet is an epitome of the Compositae flora of all China.
In the Compositae flora of Tibet, there are only 5 large genera each containing 30
species or more. They are Aster, Artemisia, Senecio, Saussurea and Cremanthodium. And
5 genera each containing 10—29 species. They are Erigeron, Anaphalis, Leontopodium,
Ajania, Ligularia and Taraxacum. In addition, there are 77 small genera, namely 87%
of the total of Compositae genera in Tibet, each comprising 1—9 species, such as Aja-niopsis, Cavea and Vernonia, etc.
2) The constituents of Compositae flora in Tibet is very closely related to those of
Sichuan-Yunnan provinces with 59 genera and 250 species in common. Such a situation
is evidently brought about by the geographycal proximity in which the Hengtuang Shan
Range links southeastern and eastern Tibet with northern and northwestern Sichuan-
Ynnnan. With India the Tibetan Compositae have 59 genera and 132 species in common,
also showing close floristic relationships between the two regions. Apparently the floris-
tic exchange of Compositae between Tibet and India is realized by way of the mountain
range of the Himalayas. The mountain range of the Himalayas, including the parallel
ranges, plays a important role as a bridge hereby some members of the Compositae of
western or northern Central Asia and of the northern Africa or of western Asia have
migrated eastwards or southeastwards as far as the southern part of Fibet and northern
part of India, or hereby some Compositae plants of eastern and southeastern Asia or
Asia Media have migrated northwestwards as the northern part of Central Asia.
Some of the species and genera in common to both Tibet and Sinjiang indicate that
this weak floristical relationship between these regions is principally realized through two
migration routes: one migration route is by way of the Himalayas including the parallel
ranges to Pamir Plataeu and Tien Shan, or vice versa. The other migration route is by
way of northern Sinjiang to Mongolia, eastern Inner Mongolia, southwards to Gansu,
Qinghai (or western Sichuan), eastern Tibet up to the Himalayas, or vice versa.
However, Tibet is not entirely situated at a migration crossroad of the floral ele-
ments. An ample amount of the data shows that Compositae flora have a particular
capability of development in Tibet. of the total number of species of Tibetan Com-
positae, 102 species and 1 genus (Ajaniopsis Shih) are endemic. Besides, 8 genera are re-
gional endemics with their range extending to its neighbourhood. The higher percentage
of endemics at specific level than at generic in Tibetan Compositae may be a result of
active speciation in response to the new enviromental conditions created by the uplifting
of the Himalayas. The flora in Tibetan Plateau as a whole appears to be of a younger
age.
3) The uprising of the Himalayas and of the Tibetan Plateau accompanied by the
ultraviolet ray radiation, the microthermal climate and the high wind pressure has, no
doubt, played a profound influence upon the speciation of the native elements of Tibetan
Compositae. The recent speciation is the main trend in the development of the Com-positae flora native in Tibet in the wake of upheaval of the plateau. 相似文献
2.
Xizang (Tibet) is rich in Leguminosae flora, comprising 41 genera and 254 species
so far known, exclusive of the commonly cultivated taxa (including 11 genera and 16
species). There are 4 endemic genera (with 8 species), 10 temperate genera (with 175
species) and 19 tropical genera (with 46 species) as well as the representatives of those
genera whose distribution centers are in East Asia-North America, Mediterranean
and Central Asia.
1. There are altogether 4 endemic genera of Leguminosae in this region. Accord-
ing to their morphological characters, systematic position and geographical distribution,
it would appear that Salweenia and Piptanthus are Tertiary paleo-endemics, while
Straceya and Cochlianths are neo-endemics. Salweenia and Piptanthus may be some
of more primitive members in the subfamily Papilionasae and their allies are largely
distributed in the southern Hemisphere. The other two genera might have been derived
from the northern temperate genus Hedysarum and the East Asian-North American
genus Apios respectively, because of their morphological resemblance. They probably
came into existanc during the uplifting of the Himalayas.
2. An analysis of temperate genera
There are twelve temperate genera of Leguminosae in the region, of which the
more important elements in composition of flora, is Astragalus, Oxytropis and Cara-
gana.
Astragalus is a cosmopolitan genus comprising 2000 species, with its center
distribution in Central Asia. 250 species, are from China so far known, in alpine zone of
Southwest and Northwest, with 70 species extending farther to the Himalayas and
Xizang Plateau.
Among them, there are 7 species (10%) common to Central Asia, 12 species (15.7%)
to Southwest China and 40 species (60%) are endemic, it indicates that the differentia-
tion of the species of the genus in the region is very active, especially in the subgenus
Pogonophace with beards in stigma. 27 species amounting to 78.5% of the total species of
the subgenus, are distributed in this region. The species in the region mainly occur in
alpine zone between altitude of 3500—300 m. above sea-level. They have developed into
a member of representative of arid and cold alpine regions.
The endemic species of Astragalus in Xizang might be formed by specialization of
the alien and native elements. It will be proved by a series of horizontal and vertical vicarism of endemic species. For example, Astragalus bomiensis and A. englerianus are
horizontal and vertical vicarism species, the former being distributed in southeast part
of Xizang and the latter in Yunnan; also A. arnoldii and A. chomutovii, the former
being an endemic on Xizang Plateau and latter in Central Asia.
The genus Oxytropis comprises 300 species which are mainly distributed in the
north temperate zone. About 100 species are from China so far known, with 40 species
extending to Himalayas and Xizang Plateau. The distribution, formation and differ-
entiation of the genus in this region are resembled to Astragalus. These two genera are
usually growing together, composing the main accompanying elements of alpine mea-
dow and steppe.
Caragana is an endemic genus in Eurasian temperate zone and one of constructive
elements of alpine bush-wood. About 100 species are from China, with 16 species in Xi-
zang. According to the elements of composition, 4 species are common to Inner Mon-
golia and Kausu, 4 species to Southwest of China, the others are endemic. This not only
indicates that the species of Caragana in Xizang is closely related to those species of
above mentioned regions, but the differentiation of the genus in the region is obviously
effected by the uplifting of Himalayas, thus leading to the formations of endemic species
reaching up to 50%.
3. An Analysis of Tropical Genera
There are 19 tropical genera in the region. They concentrate in southeast of Xizang
and southern flank of the Himalayas. All of them but Indigofera and Desmodium are
represented by a few species, especially the endemic species. Thus, it can be seen that
they are less differentiated than the temperate genera.
However, the genus Desmodium which extends from tropical southeast and northeast
Asia to Mexio is more active in differentiation than the other genera. According to Oha-
Shi,s system about the genus in 1973, the species of Desmodium distributed in Sino-Hima-
laya region mostly belong to the subgenus Dollinera and subgenus Podocarpium. The
subgenus Dollinera concentrates in both Sino-Himalaya region and Indo-China with 14
species, of which 7 species are endemic in Sino-Himalaya. They are closely related to
species of Indo-China, southern Yunnan and Assam and shows tha tthey have close con-
nections in origin and that the former might be derived from the latter.
Another subgenus extending from subtropical to temperate zone is Podocarpium.
Five out of the total eight species belonging to the subgenus are distributed in Sino-
Himalaya and three of them are endemic.
An investigation on interspecific evolutionary relationship and geographic distribu-
tion of the subgenus shows that the primary center of differentiation of Podocarpium
is in the Sino-Himalaya region.
Finally, our survey shows that owing to the uplifting of the Himalayas which has
brought about complicated geographic and climatic situations, the favorable conditions
have been provided not only for the formation of the species but also for the genus in cer-tain degree. 相似文献
3.
张志耘 《中国科学院研究生院学报》1988,26(5):394-403
The morphological characters in the genus Orobanche were evaluated from the
taxonomic point of view. The author finds that the plants of this genus are relatively similar
to each other in respect to characters of vegetative organs, fruits and seeds. But the differences
in the floral structures can be served as a basis for delimitating infrageneric taxa. The seed
coat of 18 species and pollen grains of 6 species were also examined under scanning electron
microscope (SEM). They seem to have little significance for distinguishing species.
The result supports G. Beck’s (1930) division of the genus Orobanche into 4 sections, of
which 2 occur in China, based on the characters of the inflorescence, bracteoles and calyx.
The author considers that some characters, such as anther hairy or not, upper lip of corolla
entire or not, lower lip longer or shorter than the upper one, the state of corolla-tube inflec-
tion and the hair type of filaments and plants, are important in distinguishing Chinese species.
A key to the species of Orobanche in China is given.
This genus consists of about 100 species, and is mostly confined to Eurasia, with over 60
species found in Caucasus and Middle Asia of USSR, where may be the mordern distribu-
tional centre.
Orobanche L. in China is represented by 23 species, 3 varieties and l forma. As shown in
Table 1, most species (12 species) are found in Xinjiang, which clearly shows a close floristic
relationship between this region and Middle Asia of USSR. 6 species are endemic to China,
of which 4 are confined to the Hengduan Mountains (Yangtze-Mekong-Salwin divide).
The relationships between this genus and related ones of Orobanchaceae are also discussed.
The author holds the following opinions: the genus Phelypaea Desf. should be considered as a
member of Orobanche L. Sect. Gymnocaulis G. Beck, the monotypic genus, Necranthus A.
Gilli endemic to Turkey, is allied with Orobanche L. Sect. Orobanche, the monotypic genus,
Platypholis Maxim, endemic to Bonin Is. of Japan, is far from Orobanche L. in relation and
should be regarded as a separate genus.
The 11 OTU’s, including all the sections of Orobanche L. and 7 genera of Orobanchaceae,
and 15 morphological characters were used in the numerical taxonomic treatment to test the
above-mentioned suggestions. After standardization of characters, the correlation matrices were
computerized. The correlation matrices were made to test the various clustering methods. At
last the UPGMA clustering method was chosen and its result is shown in a phenogram. The
result of numerical analysis is basically in accordance with the suggestions. 相似文献
4.
木兰科分类系统的初步研究 总被引:10,自引:0,他引:10
刘玉壶 《中国科学院研究生院学报》1984,22(2):89-109
A new system of classification of Magnoliaceae proposed. This paper deals mainly with taxonomy and phytogeography of the family Magnoliaceae on the basis of external morphology, wood anatomy and palynology. Different authors have had different ideas about the delimitation of genera of this family, their controversy being carried on through more than one hundred years (Table I). Since I have been engaged
in the work of the Flora Reipublicae Popularis Sinicae, I have accumulated a considerable amount of information and material and have investigated the living plants at their natural localities, which enable me to find out the evolutionary tendencies and primitive morphological characters of various genera of the family. According to the evolutionary tendencies of the characters and the geographical distribution of this family I propose a
new system by dividing it into two subfamilies, Magnolioideae and Liriodendroideae Law (1979), two tribes, Magnolieae and Michelieae Law, four subtribes, Manglietiinae Law, Magnoliinae, Elmerrilliinae Law and Micheliinae, and fifteen genera (Fig. 1 ), a system which is different from those by J. D. Dandy (1964-1974) and the other authors.
The recent distribution and possible survival centre of Magnoliaceae. The members of Magnoliaceae are distributed chiefly in temperate and tropical zones of the Northern Hemisphere, ——Southeast Asia and southeast North America, but a few genera and species also occur in the Malay Archipelago and Brazil of the Southern Hemisphere. Forty species of 4 genera occur in America, among which one genus (Dugendiodendron) is endemic to the continent, while about 200 species of 14 genera occur in Southeast Asia, of which 12 genera are endemic. In China there are about 110 species of 11 genera which mostly occur in Guangxi, Guangdong and Yunnan; 58 species and more than 9 genera occur in the mountainous districts of Yunnan. Moreover, one genus
(Manglietiastrum Law, 1979) and 19 species are endemic to this region. The family in discussion is much limited to or interruptedly distributed in the mountainous regions of Guangxi, Guangdong and Yunnan. The regions are found to have a great abundance of species, and the members of the relatively primitive taxa are also much more there than in the other regions of the world.
The major genera, Manglietia, Magnolia and Michelia, possess 160 out of a total of 240 species in the whole family. Talauma has 40 species, while the other eleven genera each contain only 2 to 7 species, even with one monotypic genus. These three major genera are sufficient for indicating the evolutionary tendency and geographical distribution of Magnoliaceae. It is worthwhile discussing their morphological characters and
distributional patterns as follows:
The members of Manglietia are all evergreen trees, with flowers terminal, anthers dehiscing introrsely, filaments very short and flat, ovules 4 or more per carpel. This is considered as the most primitive genus in subtribe Manglietiinae. Eighteen out of a total of 35 species of the genus are distributed in the western, southwest to southeast Yunnan. Very primitive species, such as Manglietia hookeri, M. insignis and M. mega-
phylla, M. grandis, also occur in this region. They are distributed from Yunnan eastwards to Zhejiang and Fujian through central China, south China, with only one species (Manglietia microtricha) of the genus westwards to Xizang. There are several species distributing southwards from northeast India to the Malay Archipelago (Fig. 7).
The members of Magnolia are evergreen and deciduous trees or shrubs, with flowers terminal, anthers dehiscing introrsely or laterally, ovules 2 per carpel, stipule adnate to the petiole. The genus Magnolia is the most primitive in the subtribe Magnoliinae and is the largest genus of the family Magnoliaceae. Its deciduous species are distributed from Yunnan north-eastwards to Korea and Japan (Kurile N. 46’) through Central
China, North China and westwards to Burma, the eastern Himalayas and northeast
India. The evergreen species are distributed from northeast Yunnan (China) to the
Malay Archipelago. In China there are 23 species, of which 15 seem to be very primi-
tive, e.g. Magnolia henryi, M. delavayi, M. officinalis and M. rostrata, which occur in
Guangxi, Guangdong and Yunnan.
The members of Michelia are evergreen trees or shrubs, with flowers axillary, an-
thers dehiscing laterally or sublaterally, gynoecium stipitate, carpels numerous or few.
Michelia is considered to be the most primitive in the subtribe Micheliinae, and is to
the second largest genus of the family. About 23 out of a total of 50 species of this
genus are very primitive, e.g. Michelia sphaerantha, M. lacei, M. champaca, and M.
flavidiflora, which occur in Guangdong, Guangxi and Yunnan (the distributional center
of the family under discussion) and extend eastwards to Taiwan of China, southern
Japan through central China, southwards to the Malay Archipelago through Indo-China.
westwards to Xizang of China, and south-westwards to India and Sri Lanka (Fig. 7).
The members of Magnoliaceae are concentrated in Guangxi, Guangdong and Yunnan
and radiate from there. The farther away from the centre, the less members we are
able to find, but the more advanced they are in morphology. In this old geographical
centre there are more primitive species, more endemics and more monotypic genera.
Thus it is reasonable to assume that the region of Guangxi, Guangdong and Yunnan,
China, is not only the centre of recent distribution, but also the chief survival centreof Magnoliaceae in the world. 相似文献
5.
方振富 《中国科学院研究生院学报》1987,25(4):307-313
1. The distribution of Salix species among the continents. There are about
526 species of Salix in the world, most of which are distributed in the Northern Hemisphere
with only a few species in the Southern Hemisphere. In Asia, there are about 375 species, mak-
ing up 71.29 percent of the total in the world, including 328 endemics; in Europe, about 114
species, 21.67 percent with 73 endemics; in North America, about 91 species, 17.3 percent with
71 endemics; in Africa, about 8 species, 1.5 percent, with 6 endemics. Only one species occurs
in South America. Asia, Europe and North America have 8 species in common (excluding 4
cultivated species). There are 34 common species between Asia and Europe, 14 both between
Europe and North America and between Asia and North America, 2 between Asia and Africa.
Acording to the Continental Drift Theory, the natural circumstances which promoted speciation
and protected newly originated and old species were created by the orogenic movement of the
Himalayas in the middle and late Tertiary. Besides, the air temperature was a little higher in
Asia than in Europe and North America (except its west part) and the dominant glaciers were
mountainous in Asia during the glacial epoch in the Quaternary Period. Then willows of Eu-
rope moved southwards to Asia. During the interglacial period they moved in opposite direc-
tion. Such a to-and-fro willow migration between Asia and Europe and between and North
America occurred so often that it resulted in the diversity of willow species in Asia. Those
species of willows common among the continents belong to the Arctic flora.
2. The multistaminal willows are of the primitive group in Salix. Asia has 28 species of
multistaminal willows, but Europe has only one which is also found in Asia. These 28 species
are divided into two groups, “northern type” and “southern type”, according to morphology of
the ovary. The boundary between the two forms in distribution is at 40°N. The multistami-
nal willows from south Asia, Africa and South America are very similar to each other and
may have mutually communicated between these continents in the Middle or Late Cretaceous
Period. The southern type willows in south Asia are similar to the North American multista-
minal willows but a few species. The Asian southern type willows spreaded all over the conti-
nents of Europe, Asia and North America through the communication between them before the
Quaternany Period. Nevertheless, it is possible that the willows growing in North America
immigranted through the middle America from South America. The Asian northern type mul-
tistaminal willows may have originated during the ice period.
The multistaminal willows are more closed to populars in features of sexual organs. They
are more primitive than the willows with 1-3 stamens and the most primitive ones in the ge-
nus.
3. The center of origin and development of willows Based on the above discussion it is re-
asonable to say that the region between 20°-40°N in East Asia is the center of the origin and
differentiation of multistaminal willows. It covers Southern and Southwestern China and nor-
thern Indo-China Pennisula. 相似文献
6.
我国悬钩子属植物的研究 总被引:1,自引:0,他引:1
陆玲娣 《中国科学院研究生院学报》1983,21(1):13-25
The genus Rubus is one of the largest genera in the Rosaceae, consisting of more
than 750 species in many parts of the world, of which 194 species have been recorded
in China.
In the present paper the Rubus is understood in its broad sense, including all the
blackberries, dewberries and raspberries, comprising the woody and herbaceous kinds.
So it is botanically a polymorphic, variable and very complicated group of plants.
The detailed analysis and investigation of the evolutionary trends of the main organs
in this genus have indicated the passage from shrubs to herbs in an evolutionary line,
although there is no obvious discontinuity of morphological characters in various taxa.
From a phylogenetic point of view, the Sect. Idaeobatus Focke is the most primitive
group, characterized by its shrub habit armed with sharp prickles, aciculae or setae,
stipules attached to the petioles, flowers hermaphrodite and often in terminal or axill-
ary inflorescences, very rarely solitary, druplets separated from receptacles. Whereas
the herbaceous Sect. Chamaemorus L. is the most advanced group, which is usually
unarmed, rarely with aciculae or setae, stipules free, flowers dieocious, solitary, dru-
plets adhering to the receptacles and with high chromosome numbers (2n = 56).
Basing upon the evolutionary tendency of morphological features, chromosome nu-
mbers of certain species recorded in literature and the distribution patterns of species,
a new systematic arrangement of Chinese Rubus has been suggested by the present
authors. Focke in his well-known monograph divided the species of Rubus into 12
subgenera, while in the Flora of China 8 sections of Focke were adapted, but some im-
portant revisions have been made in some taxa and Sect. Dalibarda Focke has been
reduced to Sect. Cylactis Focke. In addition, the arrangement of sections is presented
in a reverse order to those of Focke’s system. The species of Rubus in China are
classified into 8 sections with 24 subsections (tab. 3) as follows: 1. Sect. Idaeobatus,
emend. Yü et Lu(11 subsect. 83 sp.); 2. Sect. Lampobatus Focke (1 sp.); 3. Sect.
Rubus (1 sp.); 4. Sect. Malachobatus Focke, emend. Yü et Lu (13 subsect. 85 sp.); 5.
Sect. Dalibardastrus (Focke)Yü et Lu (10 sp.); 6. Sect. Chaemaebatus Focke (5 sp.);
7. Sect. Cylactis Focke, emend. Yü et Lu (8 sp.); 8. Sect. Chamaemorus Focke (1 sp.).
In respect to the geographical distribution the genus Rubus occurs throughout the
world as shown in tab. 2, particularly abundant in the Northern Hemisphere, while
the greatest concentration of species appears in North America and E. Asia. Of the
more than 750 species in the world, 470 or more species (64%) distributed in North
America. It is clearly showm that the center of distribution lies in North America at
present time. There are about 200 species recorded in E. Asia, of which the species
in China (194) amount to 97% of the total number. By analysis of the distribution
of species in China the great majority of them inhabit the southern parts of the Yangtze
River where exist the greatest number of species and endemics, especially in south-
western parts of China, namely Yunnan, Sichuan and Guizhou (tab. 3. 4.). It is in-
teresting to note that the centre of distribution of Rubus in China ranges From north-
western Yunnan to south-western Sichuan (tab. 5), where the genus also reaches its
highest morphological diversity.
In this region the characteristics of floristic elements of Rubus can be summarized
as follows: it is very rich in composition, contaning 6 sections and 94 species, about
66% of the total number of Chinese species; there are also various complex groups,
including primitive, intermediate and advanced taxa of phylogenetic importance; the
proportion of endemic plants is rather high, reaching 61 species, up to 44% of the
total endemics in China. It is noteworthy to note that the most primitive Subsect.
Thyrsidaei (Focke) Yü et Lu, consisting of 9 endemic species, distributed in southern
slopes of the Mts. Qin Ling and Taihang Shan (Fig. 4). From the above facts we may
concluded that the south-western part of China is now not only the center of distribu-
tion and differentiation of Rubus in China, but it may also be the center of origin ofthis genus. 相似文献
7.
论胡桃科植物的地理分布 总被引:1,自引:0,他引:1
路安民 《中国科学院研究生院学报》1982,20(3):257-274
The present paper aims to discuss the geog raphical distribution of the Juglandaceae
on the basis of unity of the phylogeny and the process of dispersal in the plants.
The paper is divided into the following three parts:
1. The systematic positions and the distribution patterns of nine living genera in
the family Juglandaceae (namely, Engelhardia, Oreomunnea, Alfaroa, Pterocarya, Cyclo-
carya, Juglans, Carya, Annamocarya and Platycarya) are briefly discussed. The evolu-
tional relationships between the different genera of the Juglandaceae are elucidated. The
fossil distribution and the geological date of the plant groups are reviewed. Through
the analysis for the geographical distribution of the Juglandaceous genera, the distribu-
tion patterns may be divided as follows:
A. The tropical distribution pattern
a. The genera of tropical Asia distribution: Engelhardia, Annamocarya.
b. The genera of tropical Central America distribution: Oreomunnea, Alfaroa.
B. The temperate distribution pattern
c. The genus of disjunct distribution between Western Asia and Eastern Asia:
Pterocarya.
d. The genus of disjunct distribution between Eurasia and America: Juglans.
e. The genus of disjunct distribution between Eastern Asia and North America:
Carya.
f. The genera whose distribution is confined to Eastern Asia: Cyclocarya, Platy-
carya.
2. The distribution of species
According to Takhtajan’s view point of phytochoria, the number of species in every
region are counted. It has shown clearily that the Eastern Asian Region and the Coti-
nental South-east Asian Region are most abundant in number of genera and species. Of
the 71 living species, 53 are regional endemic elements, namely 74.6% of the total species.
The author is of the opinion that most endemic species in Eurasia are of old endemic
nature and in America of new endimic nature. There are now 7 genera and 28 species
in China, whose south-western and central parts are most abundant in species, with Pro-
vince Yunnan being richest in genera and species.
3. Discussions of the distribution patterns of the Juglandaceae
A. The centre of floristic region
B. The centre of floristic regions is determined by the following two principles: a.
A large number of species concentrate in a district, namely the centre of the majority;
b. Species of a district can reflect the main stages of the systematic evolution of the
Juglandaceae, namely the centre of diversity. It has shown clearly that the southern
part of Eastern Asian region and the northern part of Continental South-east Asian
Region (i.c. Southern China and Northern Indo-China) are the main distribution centre
of the Juglandaceae, while the southern part of Sonora Region and Caribbean Region
(i.c. South-western U.S.A., Mexico and Central America) are the secondary distribution
centre.
As far as fossil records goes, it has shown that in Tertiary period the Juglanda-
ceae were widely distributed in northern Eurasia and North America, growing not only
in Europe and the Caucasus but also as far as in Greenland and Alaska. It may be
considered that the Juglandaceae might be originated from Laurasia. According to
the analysis of distribution pattern for living primitive genus, for example, Engelhar-
dia, South-western China and Northern Indo-China may be the birthplace of the most
primitive Juglandaceous plants. It also can be seen that the primitive genera and the
primitive sections of every genus in the Juglandaceae have mostly distributed in the
tropics or subtropics. At the same time, according to the analysis of morphological cha-
racters, such as naked buds in the primitive taxa of this family, it is considered that
this character has relationship with the living conditions of their ancestors. All the
evidence seems to show that the Juglandaceae are of forest origin in the tropical moun-
tains having seasonal drying period.
B. The time of the origin
The geological times of fossil records are analyzed. It is concluded that the origin
of the Juglandaceae dates back at least as early as the Cretaceous period.
C. The routes of despersal
After the emergence of the Juglandaceous plant on earth, it had first developed and
dispersed in Southern China and Indo-China. Under conditions of the stable tempera-
ture and humidity in North Hemisphere during the period of its origin and development,
the Juglandaceous plants had rapidly developed and distributed in Eurasia and dis-
persed to North America by two routes: Europe-Greenland-North America route and
Asia-Bering Land-bridge-North America route. From Central America it later reached
South America.
D. The formaation of the modern distribution pattern and reasons for this forma-
tion.
According to the fossil records, the formation of two disjunct areas was not due to
the origin of synchronous development, nor to the parallel evolution in the two con-
tinents of Eurasia and America, nor can it be interpreted as due to result of transmis-
sive function. The modern distribution pattern has developed as a result of the tectonic
movement and of the climatic change after the Tertiary period. Because of the con-
tinental drift, the Eurasian Continent was separated from the North American Conti-
nent, it had formed a disjunction between Eurasia and North America. Especially, under
the glaciation during the Late Tertiary and Quaternary Periods, the continents in Eu-
rasia and North America were covered by ice sheet with the exception of “plant refuges”, most plants in the area were destroyed, but the southern part of Eastern Asia
remained practically intact and most of the plants including the Juglandaceae were
preserved from destruction by ice and thence became a main centre of survival in the
North Hemisphere, likewise, there is another centre of survival in the same latitude in
North America and Central America.
E. Finally, the probable evolutionary relationships of the genera of the Juglanda-ceae is presented by the dendrogram in the text. 相似文献
8.
海菜花属的分类、地理分布和系统发育 总被引:1,自引:0,他引:1
李恒 《中国科学院研究生院学报》1981,19(1):29-42
The genus Ottelia is one of the great genera of Hydrocharidaceae. About 25 spe-
cies distributed in the Palaeotropics, extending from Africa through India and SE.
Asia to Korea and Japan, Australia and New Caledonia, 1 species in Brazil; centres of
specific devolopment are found in Central Africa and SE Asia.
The present study is mainly based on the materials collected during the field ex-
plorations in the lakes of Yunnan and observations on the structure of the spathe and
flowers, the variation of leaf of the plants cultivated in Kunming Bot. Garden.
Instead of the wings of the spathe used by Dandy, by the characters such as uni-or
bisexual flowers, this genus is divided into two subgenera, which by the number of the
flowers in spathe and the number of the carpus in ovary again subdivided into 4
sections. They are as the following:
A. Subg. Ottelia. Flowers bisexual.
Sect. 1. Ottelia. Spathe with 1 flower; ovary with 6(—9) carpus.
Sect. 2. Oligolobos (Gagnep.) Dandy. Spathe with many flowers; ovary with 3 car-
pus.
B. Subg. Boottia (Wall.) Dandy. Flowers unisexual; the male spathe with 1-many
flowers, the female spathe with many flowers.
Sect. 3. Boottia. The male spathe with 1 flower; ovary with 9(—15) carpus.
Sect. 4. Xystrolobos (Gagnep.) H. Li. The female spathe with (2-) many flow-
ers; ovary with 3 or 9 carpus.
The Chinense species of ottelia is in great need for revision. All of the species in
China previousely described under Ottelia Pers, Boottia Wall., Oligolobos Gagnep, and
Xystrolobos Gagen. are here combined into 3 species. They are O. alismoides, O. cor-
data, O. acuminata with 4 variaties.
After a study of the geographic distribution and infer relation-ships among the
floristic elements it has been proved that Ottelia is certainly an ancient genus, and the
primitive types came into being and widely dispersed before the separation of Laurasia
from Gondwana.
During a considerable period of time the elements of the genus Ottelia in fresh-
water environment of different continents have been separately differentiated and evolv-
ed into more or less derived types. The structure of flowers in all of the asian species
shows the following evolutionary tendenoes: 1. In this genus the plants with unisexual
flowers have evolved from plants with bisexual flower; 2. In the groups with bisexual
or unisexual flowers the number of stamens and styles reduced to 3-merous, but the
number of flowers in spathe increased. So that the subgenus Ottelia is more primitive
than the subgenus Bottia; While in the subgenus Ottelia O. alismoides is a more primi-
tive than O. balansae and in the subgenus Boottia O. cordata is the most primitive, butO. alata seems to be the most advanced. 相似文献
9.
陈重明 《中国科学院研究生院学报》1981,19(1):136-139
The “Iconographia Plantarum” written by Wu Chi-Tseng in the Qing Dynasty
(A. D. 1848) is a classical monumental work in the literature of botany. 1714 species
of plants widely-spread all over 19 provinces of our country, especeally in Jiangxi,
Hunan and Yunnan Provinces, were described in this book.
nan and Yunnan Provinces, were described try, especeally in Jiangxi, Hunan and Yun.
In order to make the plant names coincide with the original material as possible,
the author had commented upon chinese herbal in considerable detail. Most of the
plants were illustrated after their habitual appearence in somewhat clear manner.
As a means for identifying certain species of the plant, this work has been playing an
important role in development of modern botanical science in China.
In modern taxonomical books a great deal of chinese name of plants are originated
from this book, and they were available for reference to numerous researchers both at
home and abroad. On account of inadequate observation, a lot of mistakes or misleadings
occured in this book, and often been overlooked by the later botanists. Some of the
modern authors still adhere to the work “Iconographia Plantarum”and even quoting
the erroneous statement from it, thus we have to correct.
This paper annotates and commentates the misleading items of 36 species of plant,
and calls for the attention to the future readers.
相似文献
10.
鄂西神农架地区的植被和植物区系 总被引:1,自引:0,他引:1
Shennungia is generally known as “The highest mountain in Central China”. It is
situated at latitude 31°342'N., longitude 110°35'E. in western Hupeh.
The area explored is deeply cut in all sides by five V-shaped valleys, giving the
landscape a steep topography. Its summit is about 3105 meters above the sea level, and
the relative altitude is from 1000-2000 meters.
The climate of the region is warm temperate. The differences of humidity-warmth
condition between the eastern and the western flanks are quite marked.
In western Hupeh and the adjacent area of Szechuan the rugged topography still
preserves some tracts of natural forests at higher elevations. Our vegetational survey
is confined to localities above 1500 meters. The collection of plant samples of the flora
is extended to the whole mountain from the foothill to the peak. The present article
deals with only a part of the results of our survey.
1. The vertical vegetation belts of Mt. Shennungia and relationships with other
regions: The vegetation belts on the eastern and the western flanks of the mountain
are shown in diagram 2 and 3. The comparison of the vertical vegetation zones of the
Mt. Shennungia with those of the Yülungshan in N. W. Yunnan and the eastern
Himalaya to the west and with those of Hwangshan and Central Japan to the east
is shown in table 4, It shows that the plant communities of the Mt. Shennungia are of
temperate nature, and they are more closely related to those of Hwangshan in S.
Anhwei and of Central Japan than to the eastern Himalaya.
2. Floristic composition: The generic ranges of flowering plant are relatively
distinct and stable. Various distributional patterns of genera are analysized.
1) Statistics of the genera in various distributional patterns: The total number
of genera of flowering plants in this region are 762, belonging to the following four
categories. A) tropical genera 239 (31.3%), B) temperate genera 416 (54.7%), C)
endemic genera 47 (6%), and D) comsmopolitan genera 61 (8%).
2) Endemic genera: An examination of the composition of the flora in western
Hupeh reveals that 47 endemic Chinese genera occur in this mountain of which 24 are
monotypic genera, 20 oligotypic and 2 multitypic as shown in Table 4. The arborescent
genera are nearly all deciduous. They are of temperate nature.
3) Temperate genera: There are 416 genera in wastern Hupeh. They are
subdivides into the following three groups according to their distributional patterns:
A) The north temperate genera: There are 159 genera belonging to 62 families in
western Hupeh. B) Eastern Asian genera: There are 117 genera belonging to 69
families in western Hupeh. Among them 22 are common to the western Szechuan,
adjacent regions of Yunnan and the Eastern Himalaya. The remaining 95 genera are
commom to both eastern China and Japan. C) The Eastern Asian-eastern North-
American genera: Of the total 762 genera known in western Hupeh, 64 are disjunc-
tively distributed in both eastern Asia and eastern North-America.
4) The tropical genera: Of the 762 genera of the flowering plant of western
Hupeh, 239 (31%) are of tropical nature.
Finally, our survey shows: 1. Many of the primitive temperate genera and ende-
mic relicts concentrate in western Hupeh and the adjacent region of Szechuan indica-
ting that it might be one of refuges of tertiary flora. Moreover, it might also be one
of the most important regions of differentiation, development and distribution of tem-
perature flora. 2. The vegetation of this region is not only of temperate nature, but
also of a transitional nature. 3. According to an analysis of the flora and a compari-
son of the vertical distribution of the vegetation of Yülungshan and Eastern Himalaya
to the west with Hwangshan and Central Japan to the east, the floristic affinity of
western Hupeh is more closely related to eastern China and Central Japan rather than
to the Eastern Himalaya, and phytogeographically this region is intermediate between
the Sino-Himalayan and the Sino-Japanese patterns. However, the problem of phyto-
geography of western Hupeh and the adjacent region of Szechuan is a complicated
one requiring further study.
相似文献
11.
应俊生 《中国科学院研究生院学报》1994,32(5):389-410
秦岭是中国长江和黄河两大水系的分水岭,位于北纬32°5′至34°45′;东经104°30′至115°52′,最高峰达3767m。该山区是我国温带植物区系最丰实的地区之一,约有种子植物3124种,隶属于158科,892属。包括热带属220属,温带属563属,和中国特有属39属。根据该山区植物区系中各大科、主要植物群落优势种和组成种类的温带性质以及温带属在整个植物区系中的主导地位。该山区的植物区系和植被具有明显的温带性特点。特有种和非特有种的分析结果表明,该山区植物区系的特点还表现在高度特有性和以中国-日本森林植物区系为主体方面。 根据古植物学资料分析,秦岭地区植物区系的起源时间不会晚于晚白垩纪;植物群落的主要成份可能以原地生长的种类为主;秦岭及其邻近古老山区,不仅对自身的植物区系和植被具有较大的发生意义,而且对东亚植物区系具有始生性质。 相似文献
12.
13.
选择我国亚热带区域76个地区(不包括行政单元)的植物区系调查和研究资料,采用主成分分析
(PCA)、TWINSPAN聚类和样带梯度分析方法,研究了我国亚热带区域植物区系地理成分的构成特征和
各成分之间的关系;各研究区域间植物区系地理成分构成的相关性和空间分异特征;并分析了贯穿我国亚热带区域的东经108.5°~111.5°和北纬29°~31°两条样带的几种区系地理成分的分布梯度。结果表明:①15种区系地理成分可以归为热带成分、温带成分、干旱区成分和东亚中心成分4组;我国亚热带各地区区系地理成分中泛热带和北温带分布的比例最高;干旱区分布诸类比例极小;东亚中心成分的比例突出,特有性强;②样点的PCA排序明显反映了北、中、南亚热带在区系地理成分构成上的差异,但亚热带东西部的差异不明显;③样带分析清晰地反映了热带、温带成分之比的梯度格局;川东鄂西地区特有分布中心的地位和世界分布与特有成分相反的分布格局。 相似文献
14.
台湾位于欧亚大陆东南缘的海洋中,地处热带的北部和亚热带的南部,约为21°45′~15°56′N,119°18′~124°34′E,是中国最大的岛屿。它是受季风气候强烈影响的地区之一,热量丰富,雨量充沛,干湿季明显。具有一个非常丰富的岛屿和山区植物区系。就其种子植物而言,约有186科,1201属,3656 种,包括热带属742属,温带属346属。根据台湾植物区系中各大科、主要植物群落优势种和中国特有种的地理分布以及热带属在整个植物区系中的主导地位,台湾地区的植物区系主体具有明显的亚热带性质。中国台湾本地特有种十分丰富,其比例远高于中国特有种的比例。这似乎表明台湾植物区系是一个古老区系在多次地质事件侵袭后又起活化的历史演变的结果。新老成分并存、共同发展是台湾植物区系的重要特点。通过台湾全部属和非特有种在周边地区地理分布的分析,中国台湾植物区系与中国大陆的关系最为密切,是东亚植物区系的重要组成部分,因此在植物分区上应属于泛北极植物区的东亚植物区系。 相似文献
15.
太白山位居秦岭的中段,为秦岭山脉第一高峰。该山区是我国温带植物区系最丰富的地区之一,
约有种子植物1782种,隶属于125科,657属,包括热带属130属,温带属436属和特有属24属。该地
区20个较大科的种数,约占其全部植物区系的66.6%,其中特有种653种。本文对这些大科的性质及
其在植物区系和植被中的作用进行了分析讨论。在分析了全部属和植物群落优势种的地理分布的基础
上,着重讨论了该地区与其他九个山区的植物区系关系。
本文对植被垂直带划分;主要植物群落的基本特点和区系相似性;物种多样性与海拔高度变化的关
系以及生活型谱与不同坡向和海拔变化的关系等作了具体分析。对该地区主要植物群落的形成时期以
及对秦岭地区植物区系的过渡性质提出了自己的看法。从植物学角度出发,秦岭地区作为亚热带和温带
植物的分界线的提法与该地区的植物区系和植物群落性质以及水热条件情况不相符合 相似文献
16.
郎楷永 《中国科学院研究生院学报》1985,23(6):418-428
The Gongga Mountain Region, located on the eastern fringe of Qinghai-
Xizang Plateau and at the north-eastern end of Hengduan Mountains, is one of the
well-known large mountain areas in Sichuan Province. There are forty five high peaks
with the elavation of 6000 m or over in the area, among which the Gongga Mountain
is the highest one, with its summit being at the altitude of 7556 m, whereas the Dadu
River Valley in the eastern part of the area is only 1150 m above sea level; The rela-
tive height in the region is thus about 6400 m.
As we know so far the orchids in Gongga Mountain Region comprise 34 genera and
75 species with 1 variety, of which 12 species are epiphytes, 59 species with 1 variety
terrestrials and 4 species saprophytes (Table 1.).
I. The geographical distribution in the Gongga Mountain Region.
The vertical distribution of the orchids in the area.
Eastern flank: There are 39 species with I variety of orchids at 1150-2300
(-2400) m alt. in the subtropical evergreen broadleaf forest zone, of which 12 species,
such as Bulbophyllum andersonii, Dendrobium hancockii, Otochilus porrecta, etc., are
epiphytic (including a semiepiphyte, Pleione bulbocodioides), 25 species with 1 variety,
such as Bletilla formosana, Calanthe davidii, Cypripedium henryi, etc., are terrestrial,
and 2 species are saprophytic (i.e. Gastrodia elata and Neottia listeroides), the upper
limit of the real epiphytic orchids is 1800 m alt. At the altitude 2300(-2400)-3600
(-3800) m, in the coniferous forest zone, found are 23 species, including 20 terrestrial
species with 1 variety, such as Amitostigma gonggashanicum, Calanthe fimbriata, Coe-
loglossum viride, etc., 2 saprophytic species (Neottia acuminata and Risleya atropurpu-
rea) and one semiepiphytic species. There are only 2 terrestrial species (i.e. Cypripe-
dium tibeticum and Orchis chusua) at 3600(-3800)-4000m alt., in alpine shrub zone,
the upper limit of the terrestrial orchids being 4000 m alt. At 4000-4400 m alt., in
alpine meadow zone no orchids have so far been found.
Western flank: There are 14 species of orchids, such as Habenaria limprichtii,
Hemipilia flabellata, Satyrium ciliatum, etc., at 2300-2800 m alt., in the river valley
shrub zone, and they all belong to the terrestrial orchids. At the altitude of 2800-
3800m, in the coniferous forest zone found are 19 species of orchids, among which 18
species are terrestrial (such as Amitostigma monanthum, Cephalanthera longifolia, Pe-
ristylus coeloceras, etc.) and one is saprophytic. There are only 4 species of terrestrial
orchids (i.e. Cypripedium tibeticum, Gymnadenia orchidis, Orchis chusua and O. dian-
tha) at 3800-4800 m alt., in the alpine shrub-meadow zone, the upper limit of vertical
distribution of terrestrial orchids being 4400 m alt. Twelve species here are common
to the eastern flank.
II. The floristic composition of orchids in Gongga Mountain Region.
1. Twenty three species, belonging to 13 genera, are widespread in Whole East-
Asian Region.
2. Eight species, belonging to 8 genera, are the elements of the Sino-Japanese
Subregion.
3. Forty one species with 1 variety, belonging to 24 genera, are the elements of
the Sino-Himalayan Subregion, more than five times the elements of the Sino-Japanese
Subregion.
4. The floristic features of the orchids in the area.
(1) The floristic elements are mainly temperate and subtropical ones.
(2) The life form is mostly terrestrial.
(3) The species endemic to China are prolific (35 species with 1 variety, belong-
ing to 18 genera, are endemic to China, and 26 species with 1 variety are distributed
in south-western China and its adjacent region; Amitostigma gonggashanicum is endemic
to the area).
The floristic composition of orchids in the area is characterized by the dominance
of terrestrial species and temperate and subtropical East-Asian elements, though with
a few Indo-African tropical elements (such as genus Satyrium (1 species) ).
In conclusion, it may be considered that the species of orchids are abundant and
floristic elements are comparatively complex in the Gongga Mountain Region. 相似文献
17.
为探讨山地植物区系构成特征及其垂直梯度的生态意义,根据对三峡大老岭地区植被垂直样带
调查获得的植物区系资料,分析了该地区植物区系成分构成的基本特征及其随海拔梯度的变化趋势,寻找了区系平衡点的位置;并利用聚类方法分析了山地气候垂直分异对区系成分构成的影响。结果表明:①大老岭植物区系具有温带性质,但仍反映了与热带区系的历史联系,有强烈的区域性;②属的分布区类型可归为热带分布、温带分布、地中海—中亚中心和东亚中心4组,各组区系成分的垂直梯度特征不同;热带、亚热带成分与温带成分的平衡点大致位于海拔650m;③区系成分构成和属的物种数量构成的聚类分析结果一致显示了植物区系构成与山地气候和植被垂直带相对应的格局。 相似文献